原著論文?総説（査読有）

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• 133. Nakamura K, Takada H, Koike S (2025) Sex and age differences in habitat use priorities of sika deer (Cervus nippon) in highly seasonal habitats of Mount Fuji. Behaviour
• 132. Inagaki A, Allen ML, Koike S (2025) Integration of specific metrics can promote the understanding of scavenging dynamics. Frontiers in Ecology and Evolution 13: 1611982?
• 131. Koike S (2025) Variation in the Asiatic black bear Ursus thibetanus diet: a review. Mammal Study 50: ?
• 130. Shimozuru M, Nakamura S, Yamazaki J, Yanagawa Y, Tamatani H,Kuroe M, Yamazaki K, Koike S, Goto Y, Naganuma T, Tochigi K, Inagaki A, Takekoshi N, Baek S, Sato N, Honda Y, Tsubota T, Ito H (2025) Epigenetic clock in bears: A simple cost-effective blood DNA methylation-based age estimation method applicable to multiple bear species. Ecology and Evolution 15: e71424. ?
• 129. Yamazaki K, Ishizuka M, Nakayama S, Banda N, Koike S, Goto Y, Yamazaki K, Nemoto Y, Fuchs B, Arnemo MJ (2025) Lead Concentration in Blood of Asian Black Bears in Two Areas of Central Japan. Science of the Total Environment 980: 179555.
• 128. Baek S, Zedrosser A, Yamazaki K, Goto Y, Takekoshi N, Koike S (2025) Risky behavior differs between sex and season in a landscape fragmented by roads. Journal of Zoology?
• ＞プレスリリース「道路は危険？それでも渡るクマの理由」
  
• 127. Baek S, Amano T, Akasaka M, Koike S (2025) The range of large terrestrial mammals has expanded into human-dominated landscapes in Japan. Communications Earth & Environment 6: 29.?
• ＞プレスリリース「人間活動の撤退は野生動物の繁栄を促進する ―耕作放棄地の増加と温暖化が分布域を拡大」
  
• 126. Fukasawa K, Morosawa T, Nakashima Y, Takagi S, Yokoyama T, Ando M, Iijima H, Saito MU, Kumada N, Tochigi K, Yoshioka A, Funatsu S, Koike S, Uno H, Enomoto T, McShea W, Kays R. (2025) Snapshot Japan 2023: the first camera trap dataset under a globally standardized protocol in Japan. Biodiversity Data Journal 13: e1411168. ?
• 125. Tsunamoto Y, Koike S, Tayasu I, Masaki T, Kato S, Kikuchi S, Nagamitsu T, Haraguchi T, Naganuma T, Naoe S (2025) Fruiting phenology affects the direction of vertical seed dispersal by mammals and birds across mountain ranges. Oecologia 207: 24.
• 124. 吉竹　歩?山田雄作?Baek S?小池伸介（2025）GPS首輪に搭載されたActivity sensorを用いたニホンカモシカ（Capricornis crispus）の活動および非活動状態の区分の試み　―野生1個体を用いた事例―．哺乳類科学
• 123. Yasuda K, Kato D, Naoe S, Amanoa T, Yoshikawa T, Tochigi K, Koike S (2025) Does forest loss and fragmentation reduce woodpecker-associated ecosystem functions?. Global Ecology and Conservation 57: e03366. ?
• 122. Inagaki A, Sugimoto Y, Allen ML, Koike S (2024) Documentation of an Asiatic black bear preying on a living sika deer caught in a leg-hold snare trap. Ursus 35: e24.?
• 121. Sato K, Goto Y, Koike S (2025) Epizoochorous seed dispersal by two Phasianidae birds: Green pheasant (Phasianus versicolor) and Chinese bamboo partridge (Bambusicola thoracicus). Ecological Research 40: 37-43.
• 120. Takekoshi N, Fujitani A, Ohnishi N, Kozakai C, Koike S, Yamazaki K (2024) Factors affecting body weight fluctuation in free-ranging Asian black bears. Ursus35: e21. ?
• 119. Maeda N, Baek S, Tochigi K, Naganuma T, Koike S (2024) Feeding habits of Japanese serow in a warm-temperate, low-elevation, evergreen broad-leaved shrub-dominated forest. Mammal Study 49: 273-287.
• 118. Kazama K, Fujita K, Shinoda Y, Koike S (2024) Sika deer trajectory prediction considering environmental factors by timeseries transformer-based architecture. Expert Systems with Applications 250: 123630. ?
• ＞プレスリリース「周辺の環境情報を適切に考慮したニホンジカの移動経路予測を行うAI技術を開発」
  
• 117. Taniwaki T, Tamura A, Sashimura N, Naruse M, Tochigi K, Komine H, Koike S (2024) Indirect effects of low-impact thinning on insect communities in forest floor of coniferous plantations. Journal of Forest Research 29: 316-325.
• 116. Yasuda K, Taniwaki T, Amano T, Koike S (2024) Responses of a bird community to sporadic outbreaks of woody herbivorous insects in a temperate beech forest in Japan. Forest Ecology and Management 560: 121824.?
• ＞プレスリリース「繁殖期の鳥は大発生する昆虫の天敵になりうるか？ ?ハバチの大発生に同調する鳥類を特定?」
  
• 115. Dawson BM, Evans MJ, Barton PS, Soga M, Tochigi K, Koike S (2024) Drastic changes in ground-dwelling beetle communities following high-intensity deer culling: Insights from an island ecosystem. Environmental Entomology 53: 223-229.?
• 114. Inagaki A, Allen M, Maruyama T, Tochigi K, Koike S (2023) Evidence for interspecific modulation of carcass consumption among facultative scavengers in Asian temperate forest. Food Webs 37: e00324
• 113. Baek S, Shimazaki A, Zedrosser A, Naganuma T, Kozakai C, Yamazaki K, Koike S (2023) Response to human-modified landscape of an apex consumer: Sex- and season-related variations in road selection and movement pattern. Global Ecology and Conservation 46: e02603.?
• 112.Tochigi K, Steyaert S, Fukasawa K, Kuroe M, Anezaki T, Naganuma T, Kozakai C, Inagaki A, Yamazaki K, Koike S (2023) Demographic parameters of Asian black bears in central Japan. Mammal Study 48: 231-144.?
• ＞プレスリリース「メスの野生ツキノワグマの一生を探る ～個体群レベルの繁殖と死亡を定量的に評価～」
  ＞TECH+での紹介
• 111.Sato K, Goto Y, Koike S (2023) Seed attachment by epizoochory depends on animal fur, body height, and plant phenology. Acta Oecologia 119: 103914.
• ＞プレスリリース「ひっつき虫は誰が運ぶ？～動物に付着する種子の量に影響する要因の解明～」
  ＞ナゾロジーでの紹介
  ＞TECH+での紹介
  ＞大学ジャーナルでの紹介
• 110.Tanaka M, Baek S, Tochigi K, Naganuma T, Inagaki A, Dewi BS, Koike S (2023) Conditions affecting ant nesting in stumps in a temperate coniferous planted forest. Forest Ecology and Management 537: 120976.
• 109.Hiruma M, Takada H, Washida A, Koike S (2023) Dietary partitioning and competition between sika deer and Japanese serows in high elevation habitats. Mammal Research 68: 305-315.
• ＞プレスリリース「シカとカモシカの食物の分けあいと取りあい競争～生息環境によって変わる種間関係～」
  
• 108.Osugi S, Baek S, Naganuma T, Tochigi K, Allen M, Koike S (2022) The effect of decreasing human activity from COVID-19 on the foraging of fallen-fruit by omnivores. Ecology and Evolution 12: e9657. ?
• ＞プレスリリース「タヌキ?アナグマの食事スタイルは人間活動の影響を受けていた？～COVID-19がもたらした都市の野生動物の行動変化～」
  ＞ナゾロジーでの紹介
  ＞大学ジャーナルでの紹介
• 107.Takayama K, Ohnishi N, Zedrosser A, Anezaki T, Tochigi K, Inagaki A,　Naganauma T, Yamazaki K, Koike S (2023) Timing and distance of natal dispersal in Asian black bears. Journal of Mammalogy 104: 265-278.?
• ＞プレスリリース「ツキノワグマの独り立ち～子グマが出生地を離れる時期と距離を解明～」
  
• 106.Tochigi K, Steyaert S, Naganuma T, Yamazaki K, Koike S (2022) Differentiation and seasonality in suitable microsites of seed dispersal by an assemblage of omnivorous mammals. Global Ecology and Conservation 40: e02335.?
• ＞プレスリリース「果実を食べる哺乳類はどこにタネを運んでいる？～季節によって変わる種子の運び手の役割～」
  
• 105.Tezuka S, Tanaka M, Naganuma T, Tochigi K, Inagaki A, Myojo H, Yamazaki K, Allen M, Koike S (2023) Comparing information derived on food habits of a terrestrial carnivore between animal-borne video systems and fecal analyses methods. Journal of Mammalogy 104: 184-193.?
• ＞プレスリリース「クマ目線でとらえたクマの食生活～首輪に内蔵したカメラでツキノワグマの食生活を撮影～ 」
  ＞ナゾロジーでの紹介
  ＞大学ジャーナルでの紹介
• 104.Inagaki A, Allen M, Maruyama T, Yamazaki K, Tochigi K, Naganuma T, Koike S (2022) Carcass detection and consumption by facultative scavengers in forest ecosystem highlights the value of their ecosystem services. Scientific Reports 12: 16451.?
• ＞プレスリリース「森の中のシカ死体、だれが最初に見つけて食べている？～動物によるシカ死体の発見と消失のパターンを解明～」
  ＞ナゾロジーでの紹介
  ＞大学ジャーナルでの紹介
• 103.Koike S, Tochigi K, Yamazaki K (2023) Are seeds of trees with higher fruit production dispersed farther by frugivorous mammals? Journal of Forest Research 28: 64-72.?
• ＞プレスリリース「実りの豊かな木ほど遠くまでタネを散布する～哺乳類の種子散布者としての役割を定量的に評価～」
• 102.Hiruma M, Tochigi K, Kishimoto R, Kuroe M, Trentin EB, Koike S (2022) Long-term stability in the winter diet of the Japanese serow. Zookeys 1122: 39-51.?
• 101.Oki K, Soga M, Amano T, Koike S (2022) Abandonment of ski pistes impoverishes butterfly communities. Biodiversity and Conservation 31: 2083-2096.
• ＞プレスリリース「スキー場であり続けることがチョウにとっては大切～ゲレンデの閉鎖はチョウ類相を衰退させることを発見～」
• 100. Ando K, Yoshikawa T, Kozakai C, Yamazaki K, Naganuma T, Inagaki A, Koike S (2022) Composite brownian walks best explain the movement patterns of Asian black bears, irrespective of sex, seasonality and food availability. Ecological Research 37: 522-531.
• 99. Naganuma T, Nakashita R, Tochigi K, Zedrosser A, Kozakai C, Yamazaki K, Koike S (2022) Functional dietary response of Asian black bears to changes in sika deer density. Journal of Wildlife Management 86: e22218.
• ＞プレスリリース「シカの増減でクマの食生活はどう変わる？～個体で異なるツキノワグマのお食事変遷～ 」
  ＞TECH+での紹介
• 98. Allen M, Krofel M, Yamazaki K, Alexander EP, Koike S (2022) Cannibalism in bears. Ursus 33: e10.
• 97. Osugi S, Trenti EB, Koike S (2022) Effects of human activity on the fallen-fruit foraging behavior of Carnivora in an urban forest. Mammal Study 47: 113-123.
• ＞プレスリリース「人を避けてこっそり、すばやくお食事？～タヌキとアナグマの都市適応術～」
  ＞ナゾロジーでの紹介
  ＞サイエンスポータルでの紹介
• 96.Taniwaki T, Watanabe K, Komine H, Tochigi K, Yamane M, Koike S (2022) Response of specialist and generalist predators to nonprogressive annual fluctuations in herbivorous insect population. Biological Control 165: 104810.
• ＞プレスリリース「大発生のタイミングはお見通し？～ハバチの不規則な発生に同調する寄生バチを発見～」
• 95.Oki K, Soga M, Amano T, Koike S (2021) Power-line corridors in conifer plantations as novel habitat for butterflies. Journal of Insect Conservation 25:829-840.
• ＞プレスリリース「送電線下に広がるチョウの楽園～送電線はチョウの保全に貢献することを発見～」
  ＞サイエンスポータルでの紹介
  ＞FNNプライムオンラインでの紹介
  ＞KokaNetでの紹介
• 94.Baek S, Iwasaki T, Yamazaki K, Naganuma T, Inagaki A, Tochigi K, Allen ML, Kozakai C, Koike S (2021) Factors affecting predenning activity in Asian black bears. Mammal Study 46: 341-346.
• 93. Sebastian-Gonzalez E, Morales-Reyes Z, Botella F, Naves-Alegre L, Perez-Garcia JM, Mateo-Tomas P, Olea PP, Moleon M, Barbosa JM, Hiraldo F, Arrondo E, Donazar JA, Cortes-Avizanda A, Selva N, Lambertucci SA, Bhattacharjee A, Brewer AL, Abernethy EF, Turner KL, Beasley JC, DeVault TL, Gerke HC, Rhodes OE, Ordiz A, Wikenros C, Zimmermann B, Wabakken P, Wilmers CC, Smith JA, Kendall CJ, Ogada D, Frehner E, Allen ML, Wittmer HU, Butler JRA, du Toit JT, Margalida A, Oliva-Vidal P, Wilson D, Jerina K, Krofel M, Kostecke R, Inger R, Per E, Ayhan Y, Sanci M, Yilmazer U, Inagaki A, Koike S, Samson A, Perrig PL, Spencer E, Newsome TM, Heurich M, Anadon JD, Buechley ER, Gutierrez-Canovas C, Elbroch LM, Sanchez-Zapata JA (2021) Functional traits driving species role in the structure of terrestrial vertebrate scavenger networks. Ecology102:e03519.?
• 92. Naganuma T, Tanaka M, Teduka S, Steyaert S, Tochigi K, Inagaki A, Myojo H, Yamazaki K, Koike S (2021) Animal-borne video systems provide insight into the reproductive behavior of the Asian black bear. Ecology and Evolution 11:9181-9190.?
• ＞プレスリリース「知られざるクマたちの恋愛事情～ツキノワグマの繁殖行動の撮影に初めて成功～ 」
• 91. Tomiyasu T, Kayano M, Hazano K, Matsui M, Nemoto Y, Naganuma T, Koike K, Yamazaki K (2021) Associations between plasma testosterone levels and season, nutritional status, age, and body size in free-ranging male Asian black bears (Ursus thibetanus) in central Honshu, Japan. General and Comparative Endocrinology 309: 113794.
• 90. Ogawa Y, Tochigi K, Naganuma T, Dewi B S, Koike S (2021) Marking behavior of Asiatic black bears (Ursus thibetanus) at rub trees. Ursus : .
• 89. Tochigi K, Kato S, Kikuchi S, Naoe S, Koike S, Nagamitsu T (2021) Phenological shift along an elevational gradient and dispersal of pollen and seeds maintain a hybrid zone between two cherry tree species. Plant Species Biology 36: 230-245.?
• ＞森林総研チャンネル（動画）での紹介
• 88. Maekawa T, Ohara K, Zhang Y, Fukutomi M, Matsumoto S, Matsumura K, Shidara H, Yamazaki S, Fujisawa R, Ide K, Nagaya N, Yamazaki K, Koike S, Miyatake T, Kimura K, Ogawa H, Takahashi S, Yoda K (2020) Deep learning-assisted comparative analysis of animal trajectories with DeepHL. Nature Communications 11: 5316.?
• 87. Moustafa MAM, Sasaki A, Shimozuru M, Nakao R, Sashika M, Yamazaki K, Koike S, Tanaka J, Tamatani H, Tsubota T (2020) Molecular detection of apicomplexan protozoa in Hokkaido brown bears (Ursus arctos yesoensis) and Japanese black bears (Ursus thibetanus japonicus). Parasitology Research 119: 3739-3753.
• 86. Allen M, Wittmer H, Ingaki A, Yamazaki K, Koike S (2021) Food caching by bears: a literature review and new observations for Asiatic and American black bears. Ursus 32:e10.
• 85. Osugi S, Trenti EB, Koike S (2020) What determines the seedling viability of different tree species in raccoon dog latrines?. Acta Oecologica 106: 103604.
• 84. Sebastian-Gonzalez E, Morales-Reyes Z, Botella F, Naves-Alegre L, Perez-Garcia JM, Mateo-Tomas P, Olea PP, Moleon M, Barbosa JM, Hiraldo F, Arrondo E, Donazar JA, Cortes-Avizanda A, Selva N, Lambertucci SA, Bhattacharjee A, Brewer A, Abernethy E, Turner T, Beasley JC, DeVault TL, Gerke HC, Rhodes Jr OE, Ordiz A, Wikenros C, Zimmermann B, Wabakken P, Wilmers CC, Smith JA, Kendall CJ, Ogada D,